Selected publications...

Degueldre F., Mardulyn P., Kuhn A., Pinel A., Karaman C., Lebas C., Schifani E., Bračko G., Wagner H.C., Kiran K., Borowiec L., Passera L., Abril S., Espadaler X., Aron S. 2021. Evolutionary history of inquiline social parasitism in Plagiolepis ants. Molecular Phylogenetics and Evolution 155: 107016.
        Degueldre et al. - Molecular Phylogenetics & Evolution 21.pdf
Fournier D., Aron S. 2021. Hybridization and invasiveness in social insects — The good, the bad and the hybrid. Current Opinion in Insect Science 46: 1-9.
        Fournier & Aron-CurrentOpinion21.pdf
Kuhn A., Aron S., Hardy O. 2021. Detection of cryptic sex in automictic populations: theoretical expectations and a case study in Cataglyphis desert ants. Frontiers in Ecology and Evolution, in press.
Lecocq de Pletincx N., Dellicour S., Aron S. 2021. The evolution of ant worker polymorphism correlates with multiple social traits. Behavioral Ecology and Sociobiology 75:113.
        Lecocq de Pletincx et al. - BES21.pdf
Perez R., de Souza Araujo N., Defrance M., Aron S. 2021. Molecular adaptations to heat stress in the thermophilic ant genus Cataglyphis. Molecular Ecology, in press.
Perez et al. - Mol. Ecol. 21.pdf

Aron S., Deneubourg J.L. 2020. Colony co‐founding in ants is an active process by queens. Scientific Reports 10: 13539.
Aron & Deneubourg - SciRep20.pdf
Lecocq de Pletincx N., Aron S. 2020. Sociogenetic organization of the red honey ant (Melophorus bagoti). Insects 11: 755.
        Lecocq de Pletincx & Aron - Insects 20.pdf
Perez R., Aron S. 2020. Adaptations to thermal stress in social insects: recent advances and future directions. Biological Reviews 95: 1535-1553.
Perez & Aron-BiologicalReviews20.pdf

Aron S., Wehner R. 2019. Cataglyphis. In: Starr, C.K. (ed.), Encyclopedia of Social Insects. Springer Nature Switzerland AG.
Chérasse S., Dacquin P., Aron S. 2019. Mating triggers an up‐regulation of vitellogenin and defensin in ant queens. Journal of Comparative Physiology A 205 : 745-753.
Chérasse et al. - J.Comp.PhysiolA 19.pdf
Darras H., Kuhn A., Aron S. 2019. Evolution of hybridogenetic lineages in Spanish Cataglyphis ants. Molecular Ecology 28: 3073-3088.
Darras et al. - MolEcol19.pdf
Kuhn A., Darras H., Paknia O., Aron S. 2019. Repeated evolution of queen parthenogenesis and social hybridogenesis in Cataglyphis desert ants. Molecular Ecology 29: 549-564.
Kuhn et al - MolEcol19 copie.pdf
Lecocq de Pletincx N., Kuhn A., Aron S. 2019. Population structure and sociogenetic organisation in a species with ergatoid queens, the desert ant Ocymyrmex robustior. Insectes Sociaux 66: 389-401.
Lecocq et al.-InsectSoc19.pdf
Reiner Brodetzki T., Inbar S., Cohen P., Aron S., Privman E., Hefetz A. 2019. The interplay between incipient species and social polymorphism in the desert ant Cataglyphis. Scientific Reports 9: 9495.
Reiner Brodetzki et al. - Scientific Report 19.pdf

Chérasse S., Aron S. 2018. Impact of immune activation on stored sperm viability in ant queens. Proceedings of the Royal Society of London Series B 285: 20182248.
Chérasse & Aron - Proc. R. Soc.Lond.B 18.pdf
Dávila F., Botteaux A., Bauman D., Chérasse S., Aron S. 2018. Antibacterial activity of male and female sperm-storage organs in ants. Journal of Experimental Biology 221: jeb175158.
Davila et al - JEB18.pdf
Kuhn A., Darras H., Aron S. 2018. Phenotypic plasticity in an ant with strong caste genotype association. Biology Letters 14: 20170705.
Kuhn et al.-Biology Letters 18.pdf
Willot Q., Mardulyn P., Defrance M., Gueydan C., Aron S. 2018. Molecular chaperoning helps safeguarding mitochondrial integrity and motor functions in the Sahara silver ant Cataglyphis bombycina. Scientific Reports 8: 9220.
Willot et al. - Scientific Reports 18.pdf

Boulay R., Aron S., Cerdá X., Doums C., Graham P., Hefetz A., Monnin T. 2017. Social life in arid environments: The case study of Cataglyphis ants. Annual Review of Entomology 62: 305-321.
Boulay et al. - Annual Review of Entomology17.pdf
Chérasse S, Aron S. 2017. Measuring inotocin receptor gene expression in chronological order in ant queens. Hormones & Behavior 96: 116-121.
Chérasse & Aron - Hormones & Behavior 17.pdf
Dávila F., Aron S. 2017. Protein restriction affects sperm number but not sperm viability in male ants. Journal of Insect Physiology 100: 71-76.
Davila & Aron -JInsectPhysiology 17.pdf
Kuhn A., Bauman D., Darras H., Aron S. 2017. Sex-biased dispersal creates spatial genetic structure in a parthenogenetic ant with a dependent-lineage reproductive system. Heredity 119: 207-213.
Kuhn et al. - Heredity 17.pdf
Peeters C., Aron S. 2017. Evolutionary reduction of female dispersal in Cataglyphis desert ants. Biological Journal of the Linnean Society 122: 58-70.
Peeters & Aron - Biol Journal Linnean Society 2017.pdf
Willot Q., Gueydan C., Aron S. 2017. Proteome stability, heat hardening, and heat-shock protein expression profiles in Cataglyphis desert ants. Journal of Experimental Biology 220: 1721-1728.
Willot et al. - JExpBiol17.pdf

Aron S., Lybaert P., Baudoux C., Vandervelden M., Fournier D. 2016. Sperm production characteristics vary with level of sperm competition in Cataglyphis desert ants. Functional Ecology 30: 614-624.
Aron et al. - Funct. Ecol.16.pdf
Aron S., Mardulyn P., Leniaud L. 2016. Evolution of reproductive traits in Cataglyphis desert ants: mating frequency, queen number and thelytoky. Behavioral Ecology and Sociobiology 70: 1367-1379.
Aron et al. - Behav. Ecol. Sociobiol. 16.pdf
Eyer P.A., Leniaud L., Tinaut A., Aron S. 2016. Combined hybridization and mitochondrial capture shape complex phylogeographic patterns in hybridogenetic Cataglyphis desert ants. Molecular Phylogenetics and Evolution 105: 251-262.
Eyer et al. - MPE16.pdf
Fournier D., de Biseau J.C., De Laet S., Lenoir A., Passera L., Aron S. 2016. Social structure and genetic distance mediate nestmate recognition and aggressiveness in the facultative polygynous ant Pheidole pallidula. PLoS One 11: e0156440.
Fournier et al. - PLoS One16.PDF
Norman V., Darras H., Tranter C., Aron S., Hughes W.O.H. 2016 Cryptic lineages hybridize for worker production in the harvester ant Messor barbarus. Biology Letters 12: 20160542.
Norman et al. - BiolLetters16.pdf
Willot Q., Simonis P., Vigneron J.P., Aron S. 2016. Total internal reflection accounts for the bright color of the Saharan silver ant. PLoS One 11 (4): e0152325.
Willot et al. - PLoSOne16.pdf

Darras H., Aron S. 2015. Introgression of mitochondrial DNA among lineages in a hybridogenetic ant. Biology Letters 11: 20140971.
Darras & Aron - BiolLetters 15.pdf
Dávila F., Chérasse S., Boomsma J.J., Aron S. 2015. Ant sperm storage organs do not have phenoloxidase constitutive immune activity. Journal of Insect Physiology 78: 9-14.
Davila et al. - JInsectPhysiol 15.pdf
Darrouzet E., Gévar J., Guignard Q., Aron S. 2015. Production of early diploid males by European colonies of the invasive hornet Vespa velutina nigrithorax. PLoS One 10: e0136680.
Darrouzet et al. - PlosOne 15.pdf
Leniaud L., Pearcy M., Taheri A., Aron. 2015. Testing the genetic determination of the soldier caste in the silver ant. Insectes Sociaux 62: 517-524.
Leniaud et al. - Insectes Sociaux 15.pdf

Darras H., Kuhn A., Aron S. 2014. Genetic determination of female castes in a hybridogenetic desert ant. Journal of Evolutionary Biology 281: 20132396.
Darras et al. - JEB14.pdf
Darras H., Leniaud L., Aron S. 2014. Large-scale distribution of hybridogenetic lineages in a Spanish desert ant. Proceedings of the Royal Society of London B 281: 20132396.
Darras et al. - Proc.R. Soc.London.pdf
Jossart Q., Wattier R.A., Kastally C., Aron S., David B., De Ridder C., Rigaud T. 2014. Genetic evidence confirms polygamous mating system in a crustacean parasite with multiple hosts. PLoS One 9: e90680.
Jossart et al. - PLoSOne14.pdf
Mardulyn P., Thurin N., Piou V., Grumiau L., Aron S. 2014. Dispersal in the inquiline social parasite ant Plagiolepis xene. Insectes Sociaux 61: 197-202.
Mardulyn et al. - Insectes Sociaux 14.pdf
Pearcy M., Delescaille N., Lybaert P., Aron S. 2014. Team swimming in ant spermatozoa. Biology Letters 10: 20140308.
Pearcy et al. - Biology Letters14.pdf
Saar M., Leniaud L., Aron S., Hefetz A. 2014. At the brink of supercoloniality: genetic, behavioral and chemical assessments of population structure of the desert ant Cataglyphis niger. Frontiers in Ecology & Evolution 2  (article 13): 1-10.
Saar et al. - Frontiers in Ecology & Evolution 14.pdf

Doums C., Ruel C., Clémencet J., Fédérici P., Cournault L., Aron S. 2013. Fertile diploid males in the ant Cataglyphis cursor: a potential cost of thelytoky? Behavioral Ecology & Sociobiology 67: 1983-1993.
Doums et al. - Behavioral Ecology & Sociobiology13.pdf
Eyer P.A., Leniaud L., Darras H., Aron S. 2013. Hybridogenesis through thelytokous parthenogenesis in two Cataglyphis desert ants. Molecular Ecology 22: 947-955.
Eyer et al. - Molecular Ecology13.pdf
Jowers M.J., Leniaud L., Cerdá X., Alasaad S., Caut S., Amor F., Aron S., Boulay R. 2013. Social and population structure in the ant Cataglyphis emmae: Adaptive mating strategies and dispersal. PLoS One 8: e72941.
Jowers et al. - PLoSOne 13.pdf
Leniaud L., Pearcy M., Aron S. 2013. Sociogenetic organisation of two desert ants. Insectes Sociaux 60: 337-344.
Leniaud et al. - Insectes Sociaux13.pdf

Aron S. 2012. Primary sex ratio regulation by queens in ants (Formicidae) and other social Hymenoptera. Myrmecological News 17: 63-80.
Aron - Myrmecological News 12.pdf
Eyer P.A., Freyer J., Aron S. 2012. Genetic polyethism in the polyandrous desert ant Cataglyphis cursor. Behavioral Ecology 24: 144-151.
Behavioral Ecology-2012-Eyer-beheco-ars146.pdf
Fournier D., Tindo M. Kenne M., Mbenoun Masse P.S., Van Bossche V., De Coninck E., Aron S. 2012. Genetic structure, nestmate recognition and behaviour of two cryptic species of the invasive big-headed ant Pheidole megacephala. PLoS One 7: e31480.
Fournier et al. - PLoS One 12.pdf
Leniaud L., Darras H., Boulay R., Aron S. 2012. Social hybridogenesis in the clonal ant Cataglyphis hispanica. Current Biology 22: 1188-1193.
Leniaud et al. - Current Biology 12.pdf

Aron S., Timmermans I., Pearcy M. 2011. Ant queens adjust egg fertilization to benefit from both sexual and asexual reproduction. Biology Letters 7: 571-573.
Aron et al. - Biology Letters 11.pdf
Leniaud L., Hefetz A., Grumiau L., Aron S. 2011. Multiple mating and supercoloniality in Cataglyphis desert ants. Biological Journal of the Linnean Society 104: 866-876.
Leniaud et al. - Biological J. Linnean Soc. 11.pdf
Pearcy M., Hardy O., Aron S. 2011. Automictic parthenogenesis and rate of transition to homozygosity. Heredity 107: 187-188.
Pearcy et al. - Heredity 11.pdf
Thurin N., Aron S. 2011 No reversion to single mating in a socially parasitic ant. Journal of Evolutionary Biology 24: 1128-1134.
Thurin & Aron - J. Evolutionary Biology 11.pdf
Thurin N., sery N., Guimbretière R., Aron S. 2011. Colony kin structure and breeding system in the ant genus Plagiolepis. Molecular Ecology 20: 3251-3260.
Thurin et al. - Molecular Ecology 11.pdf

Debout G.D., Frederickson M.E., Aron S., Yu D.W. 2010. Unexplained split sex ratios in the Neotropical plant-ant, Allomerus octoarticulatus var. demerarae (Myrmicinae): a test of hypotheses. Evolution 64: 126-141.
Debout et al. - Evolution 10.pdf
Timmermans I., Grumiau L., Hefetz A., Aron S. 2010. Mating system and population structure in the desert ant Cataglyphis livida. Insectes Sociaux 37: 39-46.
Timmermans et al. - Insectes Sociaux 10.pdf

Aron S., Steinhauer N., Fournier D. 2009. Influence of queen phenotype, investment and maternity apportionment on the outcome of fights in cooperative foundations of the ant Lasius niger. Animal Behaviour 77: 1067-1074.
Aron et al. - Animal Behaviour 09.pdf
Cournault L., Aron S. 2009. Diploid males, diploid sperm production and triploid females in the ant Tapinoma erraticum. Naturwissenschaften 96: 1393-1400.
Cournault & Aron - Naturwisenschaften 09.pdf
Den Boer S.P.A., Baer B., Dreier S., Aron S., Nash D., Boomsma J.J. 2009. Prudent sperm use by leafcutter ant queens. Proceedings of the Royal Society of London B 276: 3945-3953.
den Boer et al. - Proc. R. Soc. Lond.B 09.pdf
Fournier D., Aron S. 2009. No-male’s land for an Amazonian ant. Current Biology 19: R738-R740.
Fournier & Aron - Current Biology 09.pdf
Fournier D., De Biseau J.-C., Aron S. 2009. Genetics, behaviour and chemical recognition of the invading ant Pheidole megacephala. Molecular Ecology 18: 186-199.
Fournier et al. - Molecular Ecology 09.pdf
Lenoir A., Aron S., Cerda X., Hefetz A. 2009. Cataglyphis desert ants: a good model for evolutionary biology in Darwin’s anniversary year - A review. In memoriam J. Kugler (1916-2007). Israelian Journal of Zoology 39: 1-32.
Lenoir et al. - Israelian J. Entomology 09.pdf
Pearcy M., Timmermans I., Allard D., Aron S. 2009. Multiple mating in the ant Cataglyphis cursor: testing the sperm limitation and the diploid male load hypotheses. Insectes Sociaux 56: 94-102.
Pearcy et al. - Insectes Sociaux 09.pdf
Thurin N., Aron S. 2009. Sib-mating in the ant Plagiolepis pygmaea: adaptative inbreeding ? Journal of Evolutionary Biology 22: 2481-2487.
Thurin & Aron - J. Evolutionary Biology 09.pdf

Cournault L., Aron S. 2008. Rapid determination of sperm number in ant queens by flow cytometry. Insectes Sociaux 55: 283-287.
Cournault & Aron - Insectes Sociaux 08.pdf
Fournier D., Bataille G., Timmermans I., Aron S. 2008. Genetic diversity, worker size polymorphism and division of labour in the polyandrous ant Cataglyphis cursor. Animal Behaviour 75: 151-158.
Fournier et al. - Animal Behaviour 08.pdf
Fournier D., Dubois D., Aron S. 2008. Isolation and characterization of microsatellite loci from the invasive ant Pheidole megacephala. Molecular Ecology Ressources 8: 919-922.
Fournier et al. - Molecular Ecology Ress. 08.pdf
Hardy O., Pearcy M., Aron, S. 2008. Small-scale spatial genetic structure in an ant species with sex biased dispersal. Biological Journal of the Linnean Society 93: 465-473.
Hardy et al. - Biological J. Linnean Soc. 08.pdf
Kapfer G., Rigot T., Holsbeek L., Aron S. 2008. Roost and hunting site fidelity of female and juvenile Daubenton’s bat Myotis daubentonii (Kuhl, 1871) (Chiroptera: Vespertilionidae). Mammalian Biology 73: 267-275.
Kapfer et al. - Mammalian Biology 07.pdf
Thurin N., Aron S. 2008. Seasonal nestmate recognition in the polydomous ant Plagiolepis pygmaea. Animal Behaviour 75: 1023-1030.
Thurin & Aron - Animal Behaviour 08.pdf
Timmermans I., Hefetz A., Fournier D., Aron S. 2008. Population genetic structure, worker reproduction and thelytokous parthenogenesis in the desert ant Cataglyphis sabulosa. Heredity 101: 490-498.
Timmermans et al. - Heredity 08.pdf

Kapfer G., Aron S. 2007. Temporal variation in flight activity, foraging activity and social interactions by rats around a sub-urban pond. Lutra 50: 9-18.
Trontti K., Thurin N., Sundström L., Aron S. 2007. Mating for convenience or genetic diversity? Mating patterns in the polygynous ant Plagiolepis pygmaea. Behavioral Ecology 18: 298-303.
Trontti et al. - Behavioral Ecology 07.pdf

Pearcy M., Aron S. 2006. Local resource competition and sex ratio in the ant Cataglyphis cursor. Behavioral Ecology 17: 569-574.
Pearcy & Aron - Behavioral Ecology 06.pdf
Pearcy M., Hardy O., Aron S. 2006. Thelytokous parthenogenesis and its consequences on inbreeding in an ant. Heredity 96: 377-382.
Pearcy et al. - Heredity 06.pdf
Schrempf A., Aron S., Heinze J. 2006. Sex determination and inbreeding depression in an ant with regular sib-mating. Heredity 97 :75-80.
Schrempf et al. - Heredity 06.pdf
Trontti K., Aron S., Sundström L. 2006. The genetic population structure of the ant Plagiolepis xene - implications for genetic vulnerability of obligate social parasites. Conservation Genetics 7: 241-250.
Trontti et al. - Conservation Genetics 06.pdf

Aron S., de Menten L., Van Bockstaele D.R., Blank S.M., Roisin Y. 2005. When hymenopteran males re-invented diploidy. Current Biology 15: 824-827.
Aron et al. - Current Biology 05
de Menten L., Fournier D., Brent C., Passera L., Vargo E.L., Aron S. 2005. Dual mechanism of queen influence over sex ratio in the ant Pheidole pallidula. Behavioral Ecology and Sociobiology 58: 527-533.
de Menten et al. - Behavioral Ecology & Sociobiology 05.pdf
de Menten L., Cremer S., Heinze J., Aron S. 2005. Primary sex ratio adjustment by ant queens in response to local mate competition. Animal Behaviour 69: 1031-1035.
de Menten et al. - Animal Behaviour 05.pdf
Passera L., Aron S. 2005. Les Fourmis. Comportement, Organisation Sociale, Evolution. Presses scientifiques du Conseil National de Recherches du Canada, Université de Western Ontario. ISBN 0-660-97021-X. 441 pp.
Trontti K., Aron S., Sundström L. 2005. Inbreeding and kinship in the ant Plagiolepis pygmaea. Molecular Ecology 14: 2007-2015.
Trontti et al. - Molecular Ecology 05.pdf

Aron S., Passera L., Keller L. 2004. Evolution of miniaturisation in inquiline parasitic ants: Timing of male elimination in Plagiolepis pygmaea, the host of Plagiolepis xene. Insectes Sociaux 51: 395-399.
Aron et al. - Insectes Sociaux 04
de Biseau J.C., Passera L., Daloze D., Aron S. 2004. Ovarian activity correlates with extreme changes in cuticular hydrocarbon profile in the highly polygynous ant, Linepithema humile. Journal of Insect Physiology 50: 585-593.
de Biseau et al. - J. Insect Physiology 04.pdf
Fournier D., Aron S., Keller L. 2004. Significant reproductive skew in the facultatively polygynous ant Pheidole pallidula. Molecular Ecology 13: 203-210.
Fournier et al. - Molecular Ecology 04.pdf
Helms K.R., Fournier D., Keller L., Passera L., Aron S. 2004. Colony sex ratios in the facultatively polygynous ant Pheidole pallidula: a reanalysis with new data. Evolution 58: 1141-1142.
Helms et al. - Evolution 04.pdf
Pearcy M., Aron S., Doums C., Keller L. 2004. Conditional use of sex and parthenogenesis for worker and queen production in ants. Science 306: 1780-1783.
Pearcy et al. - Science 04.pdf
Pearcy M., Clémencet J., Chameron S., Aron S., Doums C. 2004. Characterization of nuclear DNA microsatellite markers in the ant Cataglyphis cursor. Molecular Ecology Notes 4: 642-644.
Pearcy et al. - Molecular Ecology Notes 04.pdf

Aron S., de Menten L., Van Bockstaele D.R. 2003. Brood sex ratio determination by flow cytometry in ants. Molecular Ecology Notes 3: 471-475.
Aron et al. - Molecular Ecology Notes 03.pdf
de Menten L., Niculita H., Gilbert M., Delneste D, Aron S. 2003. Fluorescence in situ Hybridisation: a new method for determining primary sex ratio in ants. Molecular Ecology 12: 1637-1648.
de Menten et al. - Molecular Ecology 03.pdf
Fournier D., Keller L., Passera L., Aron S. 2003. Colony sex ratios vary with breeding structure but not relatedness asymmetry in the facultatively polygynous ant Pheidole pallidula. Evolution 57: 1336-1342.
Fournier et al. - Evolution 03.pdf
Roisin Y., Aron S. 2003. Split sex ratio in perennial social Hymenoptera: A mixed ESS from the queens perspective ? The American Naturalist 162: 624-637.
Roisin & Aron - American Naturalist 03.pdf

Fournier D., Aron S., Milinkovitch M. 2002. Investigation of population genetic structure and mating system in the ant Pheidole pallidula. Molecular Ecology 11: 1805-1814.

Aron S. 2001. Reproductive strategy: an essential component in the success of incipient colonies of the invasive Argentine ant. Insectes Sociaux 48: 25-27. 
Aron S., Keller L., Passera L. 2001. Role of resource availability on sex, caste and reproductive allocation ratios in the Argentine ant Linepithema humile. Journal of Animal Ecology 70: 831-839. 
Leclercq S., Charles S., Daloze D., Braekman J.C., Aron S., Pasteels J.M. 2001. Absolute configuration of anabasine from Messor and Aphaenogaster ants. Journal of Chemical Ecology 27: 945-952. 
Passera L., Aron S., Vargo E.L., Keller L. 2001. Queen control of sex ratio in fire ants. Science 293: 1308-1310.
Passera L.,  Gilbert M., Aron S.  2001. Social parasitisme in ants: effects of the inquiline parasite Plagiolepis xene St. on queens distribution and workers production of its host Plagiolepis pygmaea Latr. Insectes Sociaux 48: 74-79.

Aron S. , Passera L.  2000. Les Sociétés Animales. Evolution de la Coopération et Organisation Sociale. Bruxelles: Editions De Boeck Université, pp. 336.
Aron S., Campan E., Boomsma J.J., Passera L. 1999. Social structure and split sex ratios in the ant Pheidole pallidula. Ethology, Ecology & Evolution 11: 209-227. 
Aron S. , Passera L. 1999. Mode of colony foundation influences the primary sex ratio in ants. Animal Behaviour 57: 325-329. 
Aron S. , Passera L., Keller L. 1999. Evolution of social parasitism in ants: size of sexuals, sex ratio and mechanisms of caste determination. Proceedings of the Royal Society of London B 266: 173-177.

Bonabeau E., Theraulaz G., Deneubourg J.L., Aron S., Camazine S. 1997. Self-organization in social insects. Trends in Ecology and Evolution 12: 188 - 193.

Keller L., Aron S., Passera L. 1996. Inter-nest sex ratio variation and male brood survival in the ant Pheidole pallidula. Behavioral Ecology 7: 292-298. 
Passera L., Aron S. 1996. Early sex discrimination and male brood elimination by workers of the Argentine ant. Proceedings of the Royal Society of London B 263: 1041-1046.

Aron S., Vargo E.L., Passera L. 1995. Sex-ratio determination in monogyne colonies of the fire ant Solenopsis invicta: role of the queen and the workers. Animal Behaviour 49: 749-757.
Passera L., Aron S., Bach D. 1995. Elimination of sexual brood in the Argentine ant Linepithema humile: queen effect and brood recognition. Entomologia Experimentalis & Applicata 75: 203-212. 

Aron S., Passera L., Keller L. 1994. Queen-worker conflict over sex ratio: A comparison of the primary and the secondary sex ratios in the Argentine ant, Iridomyrmex humilis. Journal of Evolutionary Biology 7: 403-418. 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